Twenty-six fossil assemblages of land and freshwater molluscs from the Pliocene to the present, were collected from locations near the town of Meade, Kansas, U.S.A. and were analyzed (along with the extant molluscan fauna of Meade County, Kansas) to look for changes in molluscan diversity through time. The fossil assemblages were analyzed for two reasons: I) to test the hypothesis that diversity (measured as taxonomic richness, dominance, turnover and habitat type) did not change through the five million years of this study (HOI); and 2) to test the hypothesis that there is no relationship between the taxonomic composition of a molluscan assemblage and the local climate (H02)' The latter hypothesis was tested by re-examining prior molluscan paleoecological studies, by analyzing the molluscan assemblages collected for this study and through ordination analysis. The results of the first analysis clearly showed that the taxonomic structure of the assemblages changed through time (HOI was falsified), but the changes appeared to be random fluctuations. Richness ranged from 7 taxa (in assemblages AGO, FAL, RYA, SPA) to 30 (C03B) with r2 = 0.237. Dominance ranged from 1.40 (SPA) to 9.66 (RTA) with r2 = 0.113. The Habitat Ratio ranged from -0.43 (FAL) to 1.00 (X I E) with r2 = 0.029 and Turnover ranged from 0.17 (AGO, F AL) to 1.00 (RNT), when each assemblage was compared to RNT (r2 = 0.024). No discernable long-tern trend in taxonomic diversity or community composition was observed. The results of the second analysis indicate that molluscs, as a group, are not as useful as climate indicators as previously supposed. Today, both land and freshwater mollusc species are broadly geographically distributed and thus are found in a variety of locations with different local climates and vegetation. Previously, Miller (1975, 1976) grouped molluscan species with similar environmental tolerances into four units called Climate Groups and used them to infer the past climatic conditions of a region. However, only a weak climate signal was detected (ANOY A found significant differences among the Groups for minimum temperature [p < 0.0001], maximum temperature [p < 0.0001] and annual precipitation [p = 0.0082]), after the previous results were reanalyzed using the methods of this study. The fossil molluscan assemblages collected for this study displayed no climate signal (ANOY A found no significant differences among the assemblages for minimum temperature [p = 0.0714] or precipitation [p =0.691] but did find a difference in maximum temperature
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Testing the use of molluscs to infer climate with special reference to the late Cenozoic molluscs of the Meade Basin, southwest Kansas.